Tuatara

Tuatara
Temporal range: Early Miocene – present,
Northern tuatara
(Sphenodon punctatus punctatus)

Relict (NZ TCS)
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Rhynchocephalia
Suborder: Sphenodontia
Infraorder: Eusphenodontia
Clade: Neosphenodontia
Family: Sphenodontidae
Subfamily: Sphenodontinae
Genus: Sphenodon
Gray, 1831 (conserved name)
Species:
S. punctatus
Binomial name
Sphenodon punctatus
(Gray, 1842) (conserved name)
Native range (New Zealand)
Current distribution of tuatara (in black): Circles represent the North Island tuatara, and squares the Brothers Island tuatara. Symbols may represent up to seven islands.
Synonyms
  • Sphaenodon
    Gray, 1831 (rejected name)
  • Hatteria
    Gray, 1842 (rejected name)
  • Rhynchocephalus
    Owen, 1845 (rejected name)

The tuatara (/təˈtɑːrə/, Māori: [ˈtʉ.a.ta.ɾa]; Sphenodon punctatus) is a species of reptile endemic to New Zealand. Despite its close resemblance to lizards, it is the only extant member of a distinct lineage, the previously highly diverse order Rhynchocephalia. The name tuatara is derived from the Māori language and means "peaks on the back".

The single extant species of tuatara is the only surviving member of its order, which was highly diverse during the Mesozoic era. Rhynchocephalians first appeared in the fossil record during the Middle Triassic, around 244-241.5 million years ago, and reached worldwide distribution and peak diversity during the Jurassic, when they represented the world's dominant group of small reptiles. Rhynchocephalians declined during the Cretaceous, with their youngest records outside New Zealand dating to the Paleocene. Their closest living relatives are squamates (lizards and snakes). Tuatara are of interest for studying the evolution of reptiles.

Tuatara are greenish brown and grey, and measure up to 80 cm (31 in) from head to tail-tip and weigh up to 1.3 kg (2.9 lb) with a spiny crest along the back, especially pronounced in males. They have a second row of upper teeth on the roof of the mouth. They are able to hear, although no external ear is present, and have a number of unusual features in their skeleton compared to lizards, including gastralia in the belly and a completely fused temporal bar at the back of the skull.

Tuatara are sometimes referred to as "living fossils". This term is currently deprecated among paleontologists and evolutionary biologists. Although tuatara have preserved the morphological characteristics of their Mesozoic ancestors (240–230 million years ago), there is no evidence of a continuous fossil record to support the idea that the species has survived unchanged since that time.

The species has between five and six billion base pairs of DNA sequence, nearly twice that of humans.

The tuatara has been protected by law since 1895. Tuatara, like many of New Zealand's native animals, are threatened by habitat loss and introduced predators, such as the Polynesian rat (Rattus exulans). Tuatara were extinct on the mainland, with the remaining populations confined to 32 offshore islands, until the first North Island release into the heavily fenced and monitored Karori Wildlife Sanctuary (now named "Zealandia") in 2005. During routine maintenance work at Zealandia in late 2008, a tuatara nest was uncovered, with a hatchling found the following autumn. This is thought to be the first case of tuatara successfully breeding in the wild on New Zealand's North Island in over 200 years.